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Keil, D. J., M. A. Luckow & D. J. Pinkava. 1988. Chromosome studies in Asteraceae from the United States, Mexico, the West Indies, and South America. Amer. J. Bot. 75: 652–668.
Lane, M. A. & J. Li. 1993. Documented chromosome numbers 1993: 1. Chromosome number reports in Compositae with emphasis on tribe Astereae of the southwestern United States and Mexico. Sida 15: 539–546.